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Packard, A. S. "On the embryology of Limulus polyphemus." The American Naturalist 19 (1885): 722–727.


Palumbi, S. R. and B. A. Johnson. "A note on the influence of life-history stage on metabolic adaptation: the responses of Limulus eggs and larvae to hypoxia." In Physiology and biology of horseshoe crabs: Studies on normal and environmentally stressed animals, edited by J. Bonaventura et al., 115–124. New York: Alan R. Liss, Inc., 1982.


Patten, W. The evolution of vertebrates and their kin. Philadelphia: 1912.


Pearl, R. "On the behavior and reactions of Limulus in early stages of its development." Journal of Comparative Neurology & Psychology 14 (1904): 138–164.


Pearse, A. S. "On the ability of certain marine invertebrates to live in diluted sea water." Biological Bulletin 54 (1928): 453–458.


Pearson, F. C. and M. Weary. "The Limulus amoebocyte lysate test for endotoxin." BioScience 30 (1980): 461–464.


Penn, D. and H. J. Brockmann. "Nest site selection in the horseshoe crab, Limulus polyphemus." Biological Bulletin 187 (1994): 373–384.


Penn, D. and H. J. Brockmann. "Age-biased stranding and righting in horseshoe crabs, Limulus polyphemus." Animal Behavior 49 (1995): 1531–1539.


Petrocci, C. "Conchin gains momentum." American Fisherman 77, no.12 (1997): 26–28.


Pomerat, C. M. "Mating in Limulus polyphemus." Biological Bulletin 64, no.2 (1933): 242-252.


Price, Kent S. Jr. "Sand shrimp: cross-link in an esturaine food web." Estuarine Bulletin (a publication of the University of Delaware Marine Laboratories) 6, nos. 3 & 4 (1961): 12–14.